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Evolution of the wet fly: From drowned insect
to
'emerger' This
page
explores how wet flies have changed over the last 150 years
so that they now:
- work at various levels in the water column:
top, middle and
bottom
- mimic gasses trapped beneath the insect's
exoskeleton
which help it ascend from the bottom before hatching
- mimic the bulbous thorax which contains the
wing buds
- mimic emergence of the adult or dun from its
immature form
whilst breaking through the surface film
The sections below describe some important steps in the evolution
of the wet fly from early patterns which represent drowned
insects to modern patterns which cover each stage in an
insect's life-cycle.
The page concludes with some speculations on the reasons why
trout take artificial flies. These ideas are drawn from
Paul's earlier academic career which involved research into
the stimuli involved in controlling sequences of animal
behaviour. |
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Winged wet flies
- these old flies are still effective despite the fact
that we
now know that very few ephemeroptera hatch from nymph to winged
adult beneath
the water
surface
- often fished with the 'across and downstream' or
'downstream-swing' technique
- unclear what they represented (cripples, drowned
spinners?)
and therefore they fell out of widespread use
- but modern insights into the egg-laying behaviour of
some
female caddis (see below) may explain their effectiveness and
restore their popularity
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Winged wet fly: March Brown (tied by Jeff Serena) |

March Brown: Rithrogena germanica. Note the similarity betwen the
hatched natural and artificial flies |
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North Country
spiders
- often called 'soft hackled wet flies'
- may represent emergers, cripples or drowned spinners
- developed in northern England and southern
Scotland
(1857-1916)
- very sparsely dressed - short thin body of silk, one
or two
turns of hackle
- fished with short line upstream and a dead-drift, or
with a
downstream-swing
- remain an effective style of fly pattern
- modern revival prompted by Sylvester Neme's book
"The Soft Hackled Fly" published in 1975
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Partridge and Orange. Note thin short body made of tying silk and
sparse (1-2 turns) hackle. |
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Skues unweighted
nymphs
- introduced (1921-1939) by Skues for use on chalk
streams -
relatively clear and slow-flowing rivers
- incorporate a distinct dark coloured thorax to
represent wing
cases
- unweighted flies fished across and upstream just
beneath the
surface with minimal movement - dead-drift
- Skues targeted trout showing 'bulging' rises
to pre-emergent nymphs prior to a hatch
- can be difficult to detect the take unless water
surface is
relatively smooth
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Medium Olive Nymph (Skues) |

Ephemerella ignita: Blue Winged Olive nymph
Note the thin body and distinct thorax just behind the head of
the natural insect which is reproduced in the artificial fly
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Sawyer's weighted nymphs
- Skue's method developed further by Frank Sawyer who
introduced the weighted
Pheasant Tail nymph which was dead-drifted and also moved to provoke an
'induced
take'
- method developed for catching trout that are often
visible in
clear chalkstreams
- nymph cast upstream of trout so it sinks to trout's
level
- then rod tip is lifted - the 'induced take'
technique
The simplicity of this fly suggests to me that Sawyer may have
stumbled upon important stimuli that elicit a trout's feeding
response: upward movement and the ratio between thorax and body
size and shape |

Pheasant tail nymph, tied with copper wire (Sawyer) |
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Flymphs
- term 'flymph' describes the transitional hatching
stage between nymph (or pupa) and dun or adult insect
- introduced (1941-1979) by Hidy and Leisenring -
influenced by
Skues, called "the American Skues"
- fished throughout water column with movement e.g. the
'Leisenring lift'
- shaggy body materials trap gas bubbles - may mimic
gas
bubbles trapped beneath exoskeleton of pupa and nymph that
facilitate ascent and eclosion
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Transition
Flymph
(McGee ) |

In 1863 - when North Country writers were advocating thinly
dressed flies such as the Partridge and Orange (see above)
- the West Country author Cutcliffe advocated hackled
wingless wet flies. Note similarity in bushy body and hackle to
modern flymphs |
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Emergers
- Advances in macro photography enable us to study
insect
metamorphosis in great detail at each stage of the life
cycle
- Advances in our understanding of animal
behaviour reveal that artificial trout flies are
probably effective because they incorporate 'trigger'
elements found in the natural insect
- The law
of heterogeneous summation could explain how
incorporating
several 'triggers' into an artificial fly increases its
attractiveness to trout. In fact, the law suggests that these
artificial flies could be more attractive to trout than the
natural insects they are supposed to represent
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Mayfly dun shedding
its nymphal
shuck.
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The Sparkle Dun (Craig Mathews) . Note how well the tail
represents the nymphal shuck |
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- Bob Wyatts Deer Hair Emerger is probably the first
conscious
attempt to design a trout fly based on what ethologists /
behavioural ecologists call 'sign stimuli'. According to
Wyatt:
"Borrowing the
essential
features of Fran Better’s Haystack and Usual, Al
Caucci’s ComparaDun, and Hans van Klinken’s
Klinkhamer Special, the DHE is designed to present a strong
prey-image. It incorporates a couple of primary stimuli, or
‘triggers’: a visible wing and a sunk abdomen. While
suggesting natural aspects of the insect, these exaggerated
features ensure that the fly will be noticed - what behavioral
ecologists call a super-normal stimulus."
- This article
explores the application of ethology to the design of trout
flies
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Deer Hair Emerger (Wyatt) |

Paul relies on these ideas to explain-away his lack of fly-tying
skills!
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Online resources
- Photographs of aquatic
insects (in japanese) - essential
viewing to understand the problem of representing nature in fur
and feather
- Donald Nicolson's site is a catalog of new and old
fly patterns
- An excellent video showing how to tie a North
Country
spider pattern the Partridge and Yellow
- Mike
Weaver selects a dozen flies for trout on
westcountry rivers
and lakes
- Craig Mathews and John Juracek describe the Sparkle Dun
- Czech nymphs
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- North Country spiders / soft hackled wet flies
- Flymphs
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- "From nymph to flymph": Allen McGee's article
on history and development of the
flymph
- Tying Allen McGee's flymphs
- Bob Wyatt's Deer Hair Emerger
- Jeff Serena's tying of the March Brown
- Frank Sawyer tying the Pheasant Tail nymph.
Step-by-step
instructions here
- Oliver Edward's video is a superb lesson in tying
Sawyer's
Pheasant Tail nymph
- Rick Hafele's article on glossosoma
caddis
- Jeff Morgan's article on glossosoma caddis
- Tying and fishing the Sparkle Pupa
- Allen McGee's article on egg-laying caddis
- Martin Joergensen's article on his Black Funnel
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"The great charm of
fly-fishing is that we are
always learning." - Theodore Gordon |