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"Small
but perfectly
formed": Some lesser-known westcountry river flies
This page explores some of the lesser-known
species of
insects which can be found if you examine the stones in the
rivers of Dartmoor and South Devon.
Mention caddis and most anglers assume you are
talking about
one of the numerous species of relatively large insects that form
a protective case around their bodies from small stones, sand or
pieces of vegetation.
But we also have a much smaller species of caddis -
Glossosoma
In addition our rivers contain species of caddis that do not
construct a protective case around their bodies
(Rhyacophila and Hydropsyche), and - as any Summer visitor will
confirm - clouds of biting midges (Simulium) which are eagerly
consumed by trout as they struggle to break through the water
surface.
It is easy to underestimate the difficulty insects experience
breaking through the barrier between water and air, and
consequently how exposed they are to being picked off by hungry
trout. Gary LaFontaine points out that - in human terms - it is
equivalent to breaking through three feet of soil above our
heads! |
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Glossosoma
caddis
(popular name: saddle case caddis, igloo caddis,
turtle case caddis, little black short-horned caddis)
Often writers of angling books overlook small flies such as
midges and small caddis in favour of the larger and more
glamorous flies. As Gary LaFontaine wrote, "A sad fact of modern
fly-fishing is
that so much of the lore is geared to one insect,
mayflies.."
The rivers running off Dartmoor are home to large
numbers of the inconspicuous Glossosoma caddis which - as
this picture shows - are easy to overlook especially when seen
alongside a larger mayfly nymph.
Glossosoma are small (3-7 mm, hook size 18-22) caddis often
present in large numbers on the algae covered upper (illuminated)
surface of stones in fast flowing sections of local rivers. They
are thought to be the oldest (i.e. most primitive in evolutionary
terms) type of case-making caddis. The more familiar caddis which
make tubular cases from sand, stones or vegetaion are thought to
be a later evolutionary development.
Glossosoma larvae have to abandon their cases - rather than add
extra material - when they outgrow them. At this point they
may drift, or be swept, downstream which would make them
susceptible to predation by trout.
Adult Glossosoma females swim underwater to deposit their eggs.
This may explain the effectiveness of 'wings' on some
wet
flies. |
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- Large numbers can be present on top of rocks
- case is a small oval barnacle-like collection of
small bits
of stone
- glossosoma larvae cannot enlarge their case, instead
they
abandon their home when they have outgrown it, and drift
downstream = potential trout food
- larvae and pupae have pale creamy tan bodies with
dark brown
heads
- adult females dive underwater to lay eggs on rocks
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glossosoma - top view |

glossosoma - bottom view showing head of larva on right |
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- imitate glossosoma pupa with a Sparkle Pupa
- imitate uncased glossosoma larva with a tan Caddis
larva
- imitate egg-laying female with a Diving Caddis using
wet fly
swing, 'Leisenring lift', or 'induced-take techniques
with weighted leader
- adults are small and grey-black in colour

Diving Caddis (Gary LaFontaine)
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Sparkle Pupa (Gary LaFontaine) |

Tan Caddis larva |
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Free-living
caddis:
Rhyacophila (popular name:Sandfly), and
Hydropsyche (net-spinning caddis)
- free-living or caseless caddis; the larvae does not
construct
a case
- Rhyacophilia are green colour
- relatively large - 9-15mm in size
- 'Czech nymph' method of fishing mimics behaviour and
location of these insects
- Czech nymphing is a short-line deep-water technique
which
involves 'flicking' rather than casting a very short line
upstream and allowing the flies to dead drift downstream. This
process can be combined with an 'induced take' by lifting
the rod.
- the heavy caseless caddis fly can be combined with a
dry fly
which acts as an 'indicator' when a fish takes the
caddis- the 'New Zealand dropper' set-up.
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Rhyacophilia (caseless caddis) larvae |

Hydropsyche larvae |
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Hans van Klinken's caseless caddis
Sawyer's Killer Bug |
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Simulium
(popular
names: biting midges, black gnat, smuts)
- club-shaped larvae found attached by a sucker to
stones,
prominent mouth 'brushes'
- pupa floats to surface in a bubble of air
- midge pupae are important trout food
- responsible for 'smutting rise'
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Simulium larvae
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midge pupa

Martin Joergensen's Black Funnel |
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Online
resources
- Rick Hafele's article on Glossosoma
caddis
- Jeff Morgan's article on Glossosoma caddis
- Jeff Morgan's tying of the uncased Glossosoma
larva
- Allen McGee's article on egg-laying caddis
- Westfly article on saddle case caddis
- Martin Joergensen's article on his Black Funnel
- Rick Hafele's article on emergence behaviours
- Rick Hafele's article on micro-caddis
- Pictures of caseless caddis
(Rhyacophilia)
larvae
- Hans van Klinken's tying and fishing the caseless
caddis
- Frank Sawyer's Killer Bug
- Czech nymphs
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- William H. Amos, article 'In Praise of
Blackflies'
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| anabolia
nervosa:
this caddis attaches bits of stick to its case |
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"The great charm of
fly-fishing is that we are
always learning." - Theodore Gordon |