The survey sites were very similar to these images of the Upper Yealm
Fishery beat. The main flow types were run and riffle, with sand and pebbles/gravels dominating the substrate. The wide variety of river flies found in the River Yealm is typical of clean, fast flowing, stony rivers.
The survey found Elmis aenea a very small (2 mm) riffle beetle that is equipped with strong claws to enable it to grip in strong currents. It is a dark coloured species with very deeply ridged wing cases.It does not need to surface for air as it breathes the trapped oxygen in submerged bubbles but it does leave the water at times and can fly. If disturbed it will float to the surface.They may move downstream by drifting in the current.
Here is a large image of Elmis aenea, useful for fly-tyers.
Dr JM Elliott (1967) studied the food of trout in the Walla Brook a tributory of the East Dart. His results highlighted the importance of beetles in trout diet.
Mike Weaver recommends this easy-to-tie simple Black Midge